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To support comparative genomics, we filled in divergence gaps in the phylogeny by generating draft genomes for eight new species.

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Approximately 13,000 gene models developed by a combination of ab initio predictions and expert manual curation were included in the first release of the research community and the Fly Base curators have resulted in an ever improving annotation (Mc Quilton et al. However, results from early microarray expression profiling studies demonstrated that transcribed elements were unannotated (Andrews et al. A classic example is the expressed pseudogene, which can arise by gene duplication followed by degeneration of one redundant copy by random accumulation of mutations (Balakirev and Ayala 2003; Zheng et al. Functionless transcripts might also arise from transposon promoters (Emera and Wagner 2012; Hancks and Kazazian 2012).

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